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The project that became Firefox today began as an experimental branch of the Mozilla Suite called m/b (or mozilla/browser). Firefox retains the cross-platform nature of the original Mozilla browser, using the XUL user interface markup language. The use of XUL made it possible to extend the browser's capabilities through the use of extensions and themes. The development and installation processes of these add-ons raised security concerns, and with the release of Firefox 0.9, the Mozilla Foundation opened a Mozilla Update website containing "approved" themes and extensions. The use of XUL sets Firefox apart from other browsers, including other projects based on Mozilla's Gecko layout engine and most other browsers, which use interfaces native to their respective platforms (Galeon and Epiphany use GTK+, K-Meleon uses MFC, and Camino uses Cocoa). Many of these projects started before Firefox, and may have served as inspiration.

Hyatt, Ross, Hewitt and Chanial developed their browser to combat the perceived software bloat of the Mozilla Suite (codenamed, internally referred to, and continued by the community as SeaMonkey), which integrated features such as IRC, mail, news, and WYSIWYG HTML editing into one internet suite. After it was sufficiently developed, binaries for public testing appeared in September 2002 under the name Phoenix. This name carried the implication of the mythical firebird that rose triumphantly from the ashes of its dead predecessor, in this case Netscape Navigator which lost the "First browser war" to Microsoft's Internet Explorer. The name Mozilla began as the internal codename for the original 1994 Netscape Navigator browser aiming to displace NCSA Mosaic as the world's most popular web browser. The name for this would-be "Mosaic killer" was meant to evoke the building-crushing Godzilla.
Due to continuing pressure from the Firebird community, on February 9, 2004, the project was renamed again to Mozilla Firefox. The name "Firefox" (a reference to the red panda) was chosen for its similarity to "Firebird", and its uniqueness in the computing industry. To ensure that no further name changes would be necessary, the Mozilla Foundation began the process of registering Firefox as a trademark with the United States Patent and Trademark Office in December 2003. This trademark process led to a delay of several months in the release of Firefox 0.8 when the foundation discovered that Firefox had already been registered as a trademark in the UK for Charlton Company software.

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